AMERICAN COCKROACH | PERIPLANETA AMERICANA (LINNAEUS) - SEMINAR WORK

CHAPTER ONE
INTRODUCTION
The American cockroach, periplaneta americana(Linnaeus), is the largest of the common peridometic cockroaches measuring on average 4cm in length, it occurs in buildings throughout Florida especially in commercial buildings. In the Northern United States, the cockroach is mainly found in steam heat tunnels or large institutional buildings. The American cockroach is second only to the German cockroach in abundance.


DISTRIBUTION/ORIGIN OF AMERICAN COCKROACH (PERIPLANETA AMERICANA)
            Forty-seven species are included in the genus, periplaneta, none of which are endemic to the US (Bell and Adiyodi 1981). The American cockroach peripaneta americana was introduced to the United States as early as1625 (Bell and Adiyodi 1981).
            The American cockroach has spread throughout the world by commerce. The cockroach is often found residing indoors as well as outdoors. It is found mainly in basements, sewers, steam tunnels, and drainage systems (Rust et. al. 1991). This cockroach is readily found in commercial and large buildings such as restaurants, grocery stores, bakeries and where food is prepared and stored. The American cockroach is rarely found in houses, however after heavy rain infestations of the cockroach can occur in homes. The can develop to enormous numbers greater than 5,000 sometimes being found in individual sewer manholes (Rust et.al.1991).
            American cockroaches are found in moist shady areas outdoors, in yards, hollow trees, wood piles, and mulch. They are occasionally found under roof shingles and in attics. The cockroaches dwell outside but will wander indoors for food and water or during extremes in weather conditions. In Florida, areas such as trees, wood piles, garbage facilities, and accumulations of organic debris around homes provide adequate food, water and harborages for peridomestic cockroaches such as the American cockroach (Hagenbuch et al. 1988). Mass migrations of the American cockroaches are common (Ebeling 1975). The migrate by crawling or flying into structures often entering houses and apartments from sewers via the plumbing, by trees and shrubs located along-side buildings or trees with branches over hanging roofs facilitate the entry of cockroaches into the home. During the day, the American cockroach, which responds negatively to light, rests in harborages close to water pipes, sinks, baths and toilets, for example, where the microclimate is suitable for survival (Bell and Adiyodi 1981).

CHAPTER TWO

CLASSIFICATION

SCIENTIFIC CLASSIFICATION, AMERICAN COCKROACH (Periplaneta americana)
Kingdom:           Animalia
Phylum:            Arthropoda
Class:                Insecta
Order:              Blattodea
Family:             Blattidae
Genus:             Periplaneta
Species:            Periplaneta americana
             Binomial name
Periplaneta Americana (Linnaeus 1758).

CHARACTERISTICS/HABITAT

            American cockroach adults are 1 and ½ inches long (38mm). They are reddish-brown and have a yellowish margin on the body region behind the head. Immature cockroaches resemble adults except that they are wingless.
        American cockroaches generally like in moist areas, but can survive in dry areas if they have access to water. They prefer warm temperatures around 29 degrees Celsius (84Af) and do not tolerate cold temperatures. In residential areas, these cockroaches live in basements and sewers and may move outdoors into yards during warm weather. These cockroaches are common in basements, crawl spaces, cracks and crevices of porches, foundations, and walkways adjacent to buildings. They feed on a wide variety of plant and animal material. The American cockroach is a scavenger that feeds on decaying organic matter and variety of other foods. It is particularly fond of fermenting foods.

CHAPTER THREE

DESCRIPTION/REPRODUCTION

Egg: Females of the American cockroach lay their eggs in a hardened, purse-shaped egg case called an OOTHECA. About one week after mating, the female produce an ootheca and at the peak of her reproductive period, she may form about two ootheca per week (Bell and Adiyodi 1981). The females on average produce an egg case about once a month for ten months laying 16 eggs per egg case. The female deposits the ootheca near a source of food by either simply dropping it or gluing it to a surface with a secretion from her mouth. The deposited ootheca contains water sufficient for the eggs to develop without receiving additional water from the substrate (Bell and Adiyodi 1981).
            The egg case is brown when deposited, and turn black in a day or two. A typical egg case contains about 14-16 eggs. It is about 8mm long and 5mm high.
            Larva on Nymph: the nymphal stage begins when the egg hatches and ends with the emergence of the adult. The number of times an American cockroach molts varies from 6-14(Bell and Adiyodi 1981). The first instars American cockroach is white immediately after hatching then become greenish-brown. After molting instars of the cockroach nymphs are white and then become a uniformly reddish-brown with the posterior margins of the thoracic and abdominal segments being a darker color.
            Wings are not present in the nymphal stages and wig pads become noticeable in the third or fourth instars. Complete development from egg to adult is about 600days.
            The nymphs as well as the adults actively forage for food and water.
Adult: The adult American cockroach is reddish-brown in appearance with a pale-brown or yellow band around the edge of the pronotum.
The males are longer than the females because their wings extend 4mm to 8mm beyond the tip of the abdomen.  Males and females have a pair of slender, jointed cerci at the tip of the abdomen. The male cockroaches have cerci with 18 to 19 segments. The male American cockroaches have pair styli between the cerci while the female do not.

LIFE CYCLE
The American cockroach has three life stages. The egg, a variable number of nymphal instars and adult. The life cycle from egg to adult averages about 600days while the adult life span may be another 400days. The immature emerges from the egg case in about 6-8 weeks and matures in about 6-12 months. Adults can live up to one year and an adult female will produce an average of 150 young in her life time. Environmental factors such as temperature and humidity can increase or decrease the developmental time of the American cockroach. Outdoors the female shows a preference for moist, concealed oviposition site (Bell and Adiyodi 1981).

CHAPTER FOUR

MANAGEMENT

            Several hymenopteran natural enemies of the American cockroach have been found (suiter et. al. 1998). These parasitic wasps deposit their eggs in the cockroach ootheca preventing the emergence of cockroach nymphs.
            Caulking of penetrations through ground level walls, removal of rotting leaves and limiting the moist areas in and around a structure can help in reducing areas that are attractive to those cockroaches.
            Other means of management are insecticides that can be applied to basement walls, wood scraps, and other infested locations. Residual sprays can be applied inside and around perimeter of an infested structure. When insecticides and sprays are used to manage cockroach populations, they may ultimately kill off the parasitic wasps. Loose, toxic, pellet baits are extremely effective in controlling American cockroach populations.
            Insect management (1): Guide for cockroaches. Cockroaches are pests throughout the United States. They are annoying, and when abundant, they are also destructive. Cockroaches, also known as water bugs, croto bugs, or palmetto bugs, destroy food and damage fabrics, book-bindings, and other materials. When cockroaches run over food, they leave filth and may spread disease. They secrete an oily liquid that has an offensive and sickening odor that may ruin food. This odor may also be imparted to dishes that apparently clean. Excrement in the form of pellets or ink-like liquid also contributes to this nauseating odor. Some people are allergic to cockroaches and become ill. 
                         KINDS OF COCKROACHES
            The kinds of cockroaches most commonly found in and around Florida homes are the Florida wood roach, American, smoky brown, Brown, Australian, German, and Asia. The smallest cockroaches, the German, Asian and Brown banded are close to the same size and the adults are seldom more than 5/8 inches long. The larger cockroaches, the American, Australian, Brown, and the smoky brown, are 1 1/4inches to 2 inches long and are often called palmetto bugs.
            Though they are generally found outdoors, they can become an indoor problem when they migrate or are carried indoors. The largest cockroach, the Florida woods roach, will also enter dwellings from the outside or from beneath the house. Outdoor cockroaches do not survive well indoors and many times people over react to the presence of these outdoor cockroaches from the house is all that is needed for control. The Asian cockroach looks very much like a German cockroach, except it flies. Asian cockroaches are attracted to the lights in your home. They will fly to porch lights and find their way inside. Once inside, however, their survival is poor like other outdoor cockroaches.
                      DEVELOPMENT OF THE COCKROACH
            The cockroach has life stages: the egg, nymph, an adult. Cockroach eggs are deposited in groups in a leathery case or capsule called an OOTHECA. This capsule is usually dropped or glued to some surface by the female and soon as it is deposited; however, the female German cockroach carries the capsule protruding from body until the eggs are ready to hatch. There may be from 30-48 eggs in the capsule of German cockroach, but capsules of other cockroaches may have only 10-28 egg.
            The newly hatch nymphs have no wings and shed their skins (molt) several times before becoming winged adults. German and brown banded cockroaches may have several generations per year, but the outdoor species may require a year to develop from egg to adult.
 
CHARTER FIVE
                    DIET/MODE OF FEEDING
            The American cockroaches are an omnivorous and opportunistic feeder. It consumes decaying organic matter but since the cockroach is a scavenger, it will eat most anything. It prefers sweets and has been observed eating paper, boots, hair, bread, fruits, book-bindings, fish, peanuts, old rice, putrid sack, the soft part on the inside of animal hides, cloth, and dead insects (Bell and adiyodi 1981).

MEDICAL AND ECONOMIC SIGNIFICANCE

American cockroaches can become public health problem due to their association with human waste and disease, and their ability to move from sewers into homes and commercial establishments. In the United States during the summer, alleyways and yards may be over-run by these cockroaches. The cockroach is found in caves, mines, privies, latrines, cesspools, sewers, sewerage treatment plants, and dumps (Bell and adiyodi 1981).
            Their presence in these habitats is of epidemiological significance. At least 22 species of pathogenic human bacteria, virus, fungi, and protozoans, as well as 5 species of helminthic worms, have been isolated from field collected American cockroaches (Rust et. al. 1991). Cockroaches are also aesthetically displeasing because they can soil items with excrement and regurgitation.

CONCLUSION

German cockroaches are the most important pest in the indoor environment. Peridomestic cockroaches live primarily outdoors but often invade structures looking for food, warmth or moisture. The treatment measures for indoor versus outdoor cockroaches are very different, so it is extremely important that a problem cockroach population be correctly identified. Once the cockroach and its habitat have been determined, the magnitude and the location of the population need to be evaluated. These can be done by performing a thorough inspection in and round the structure and monitoring with traps.   The population information should then be used to choose treatment strategies. A combination of treatments is recommended for a complete approach to cockroach management.
            Several least toxic treatment choices are available for cockroach control, they includes: bait products (available for indoor and outdoor use) insect growth regulators (IGRs), inorganic ducts and traps. Oothecal parasitoids occur as a biological control for peridomestic cockroaches.
            However, these wasps are very sensitive to insecticides and should be protected from outdoor applications, particularly those involving pyrethroids.

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